Sphingolipids (SPL), the general name for lipids containing a sphingoid long-chain base, are one of the most chemically and functionally diverse classes of biomolecules . Sphingomyelin (SM) and glycosphingolipids (GSPL) constitute the major complex SPL in mammalian cells, both of which contain Cer as a lipid portion and phosphorylcholine or sugars, respectively, as a hydrophilic moiety. Cer, composed of a sphingoid base (SB) and an amide-linked fatty acid (FA), serves as the precursor for almost all SPL .
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